Cole and Tuilagi are joined in the England starting XV by Leicester Tigers teammates Tom Croft, Louis Deacon and Ben Youngs. Toby Flood is named among the replacements.
Martin Johnson makes three changes to his team with Delon Armitage starting on the wing in place of Mark Cueto, lock Courtney laws replacing Tom Palmer and Matt Stevens in at loosehead for prop Alex Corbisiero.
Bath Rugby flanker Lewis Moody again captains the side on the occasion of his 70th cap and fly half Jonny Wilkinson will make his 90th Test appearance for England.
England
15 Ben Foden (Northampton Saints)
14 Chris Ashton (Northampton Saints)
13 Manusamoa Tuilagi (Leicester Tigers)
12 Mike Tindall (Gloucester Rugby)
11 Delon Armitage (London Irish)
10 Jonny Wilkinson (Toulon)
9 Ben Youngs (Leicester Tigers)
1 Matt Stevens (Saracens)
2 Steve Thompson (London Wasps)
3 Dan Cole (Leicester Tigers)
4 Louis Deacon (Leicester Tigers)
5 Courtney Lawes (Northampton Saints)
6 Tom Croft (Leicester Tigers)
7 Lewis Moody (Bath Rugby, capt)
8 James Haskell (Ricoh Black Rams)
Replacements
16 Dylan Hartley (Northampton Saints)
17 Alex Corbisiero (London Irish)
18 Tom Palmer (Stade Francais)
19 Nick Easter (Harlequins)
20 Richard Wigglesworth (Saracens)
21 Toby Flood (Leicester Tigers)
22 Matt Banahan (Bath Rugby)
