Cole and Croft replace David Wilson (Bath Rugby) and James Haskell (Stade Francais) respectively and join Leicester Tigers teammates Toby Flood and Ben Youngs in the starting line-up.
Johnson makes four changes to the team that beat Samoa 26-13 with captain Lewis Moody coming in for Hendre Fourie (Leeds Carnegie) and Mike Tindall (Gloucester Rugby) in for Matt Banahan (Bath Rugby).
Fourie, Wilson and Banahan are on the bench, where Simon Shaw (London Wasps) returns in place of David Attwood (Gloucester Rugby).
Moody draws level with Neil Back as England’s most-capped flanker when he makes his 66th international appearance. Although Joe Worsley is the most experienced back-row forward in the England squad, 15 of his 77 caps have come at No.8.
ENGLAND TEAM
15 Ben Foden (Northampton Saints)
14 Chris Ashton (Northampton Saints)
13 Mike Tindall (Gloucester Rugby)
12 Shontayne Hape (Bath Rugby)
11 Mark Cueto (Sale Sharks)
10 Toby Flood (Leicester Tigers)
9 Ben Youngs (Leicester Tigers)
1 Andrew Sheridan (Sale Sharks)
2 Dylan Hartley (Northampton Saints)
3 Dan Cole (Leicester Tigers)
4 Courtney Lawes (Northampton Saints)
5 Tom Palmer (Stade Francais)
6 Tom Croft (Leicester Tigers)
7 Lewis Moody (Bath Rugby, captain)
8 Nick Easter (Harlequins)
REPLACEMENTS
16 Steve Thompson (Leeds Carnegie)
17 David Wilson (Bath Rugby)
18 Simon Shaw (London Wasps)
19 Hendre Fourie (Leeds Carnegie)
20 Danny Care (Harlequins)
21 Charlie Hodgson (Sale Sharks)
22 Matt Banahan (Bath Rugby)
